The Evolution of Man — Volume 2 by Haeckel, Ernst Heinrich Philipp August, 1834-1919

We must grant, however, that in the whole stem-history of the Vertebrates the long stretch from the Gastraeads and Platodes up to the oldest Chordonia remains by far the most obscure section. We might frame another hypothesis to raise the difficulty--namely, that there was a long series of very different and totally extinct forms between the Gastraea and the Chordaea. Even in this modified chordaea-theory the six fundamental organs of the chordula would retain their great value. The medullary tube would be originally a chemical sensory organ, a dorsal olfactory tube, taking in respiratory-water and food by the neuroporus in front and conveying them by the neurenteric canal into the primitive gut. This olfactory tube would afterwards become the nervous centre, while the expanding gonads (lying to right and left of the primitive mouth) would form the coeloma. The chorda may have been originally a digestive glandular groove in the dorsal middle line of the primitive gut. The two secondary gut-openings, mouth and anus, may have arisen in various ways by change of functions. In any case, we should ascribe the same high value to the chordula as we did before to the gastrula.

In order to explain more fully the chief stages in the advance of our race, I add the hypothetical sketch of man's ancestry that I published in my Last Link [a translation by Dr. Gadow of the paper read at the International Zoological Congress at Cambridge in 1898]:--

COLUMN 1 : CHIEF STAGES. COLUMN 2 : ANCESTRAL STEM-GROUPS. COLUMN 3 : LIVING RELATIVES OF ANCESTORS.

STAGES 1 TO 5. PROTIST ANCESTORS. UNICELLULAR ORGANISMS.

1 to 2. Protophytes. : 1. Monera. Without nucleus. : Chromacea. (Chroococcus.) Phycochromacea.

1 to 2. Protophytes. : 2. Algaria. Unicellular algae. : 2. Paulotomea. Palmellacea. Eremosphaera.

3 to 5. Protozoa. : 3. Lobosa. Unicellular (amoebina) rhizopods. : 3. Amoebina. Amoeba Leucocyta.

3 to 5. Protozoa. : 4. Infusoria. Unicellular. : 4. Flagellata. Euflagellata. Zoomonades.

3 to 5. Protozoa. : 5. Blastaeades. Multicellular hollow spheres. : 5. Catallacta. Magosphaera, Volvocina, Blastula.

STAGES 6 TO 11. INVERTEBRATE METAZOA ANCESTORS.

6 to 8. Coelenteria, without anus and body-cavity. : 6. Gastraeades. With two germ-layers. : 6. Gastrula. Hydra, Olynthus, Gastremaria.

6 to 8. Coelenteria, without anus and body-cavity. : 7. Platodes I. Platodaria (without nephridia). : 7. Cryptocoela. Convoluta, Proporus.

6 to 8. Coelenteria, without anus and body-cavity. : 8. Platodes II. Platodinia (with nephridia). : 8. Rhabdocoela. Vortex, Monotus.

9 to 11. Vermalia, with anus and body-cavity. : 9. Provermalia. (Primitive Worms.) Rotatoria. : 9. Gastrotricha. Trochozoa, Trochophora.

9 to 11. Vermalia, with anus and body-cavity. : 10. Frontonia. (Rhynchelminthes.) Snout-worms. : 10. Enteropneusta. Balanoglossus, Cephalodiscus.

9 to 11. Vermalia, with anus and body-cavity. : 11. Prochordonia. Chorda-worms. : 11. Copelata. Appendicaria. Chordula-larvae.

STAGES 12 TO 15. MONORHINA ANCESTORS.

Oldest vertebrates without jaws or pairs of limbs, single nose. : 12. Acrania I. (Prospondylia.) : 12. Amphioxus larva.

Oldest vertebrates without jaws or pairs of limbs, single nose. : 13. Acrania II. More recent. : 13. Leptocardia. Amphioxus.

Oldest vertebrates without jaws or pairs of limbs, single nose. : 14. Cyclostoma I. (Archicrania.) : 14. Petromyzonta larvae.

Oldest vertebrates without jaws or pairs of limbs, single nose. : 15. Cyclostoma II. More recent. : 15. Marsipobranchia. Petromyzonta.

B. MAN'S GENEALOGICAL TREE, SECOND HALF: LATER ANCESTORS, WITH FOSSIL EVIDENCE.